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Hi Eric, Between 1931 and 1933 the Flying Squirrel used the single downtube frame. The duplex frame was reintroduced in late 1933, for the 1934 ‘season’ but the frame geometry was different from that used between late 1928 and late 1933. (The duplex frame was retained for the TT Replica only 1931-1933). So the answer to your question is late 1933 onwards, and nothing to do with twin-filler to single-filler tanks.